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RESEARCH PLAN
1. the project
Abstract
of Proposal
The objectives of this project are the discovery and extraction of
vertebrate and microvertebrate fossils from the Upper Triassic Ischigualasto
Formation (Carnian) and Los Colorados Formation (Norian) in northwest
Argentina. Fossils from this period, especially carnians, are very rare in
the rest of the world and are uniquely preserved in outcrops of the
Ischigualasto Basin. Results of discoveries made in Ischigualasto
contribute to the knowledge of the early history and evolution of several
vertebrates groups such as: archosauriforms, primitive archosaurs (including
the oldest record of primitive dinosaurs), therapsids, turtles,
sphenodontids and labyrinthodon amphibians. The top of the Los Colorados
Formation will be mapped, and several sections will be made, in order to
determine the presence of Lower Jurassic sediments lying on top of the
Formation. Several cross-sections will be made in order to find any
structural change. We will try to relocate all the findings made in the last
30 years using old bibliography, documents and surviving local guides. All
the findings will be mapped using aerial photographs and topographic maps.
They also will be located in detailed stratigraphic sections for the
analysis of biostratigraphic changes through the Upper Triassic. The
relationships between the sediments in which vertebrate remains are found
and the appearance of the skeletons will be documented for taphonomic
analysis. As part of a collaborative agreement with the University of
Arizona, data will be collected to document large-scale paleoclimate change
in the Triassic rocks of the Ischigualasto Basin.
Motivated volunteers can contribute greatly to the prospecting and
extraction of fossils. Volunteer assignments will be the same as those of
the professional staff.
RESEARCH
PLAN
1. The
Project
Our goal is
to contribute to the knowledge of the early history of dinosaurs, crocodiles
and mammals. The early Mesozoic era, and especially the Upper Triassic, was
a critical period in the evolution of life on land. Most of today’s major
groups of terrestrial vertebrates (mammals, crocodiles, turtles, lizards,
frogs, salamanders, etc.) originated, and dinosaurs and pterosaurs rose to
prominence during this period. In recent years this period in the earth’s
history has received a great deal of attention, and, in particular, it is
now believed that the small vertebrates that lived in the shadows of the
first dinosaurs can tell us a great deal about the evolution of what might
be termed modern terrestrial ecosystems.
The early
part of the Triassic was basically a continuation of the Permian, dominated
by synapsids, especially cynodonts, in the herbivore and carnivore
ecologies. The Middle Triassic was a transition between the synapsids’
dominance to the archosaurs’ dominance. In the late Triassic the synapsids’
dominance declined, and the world was full of archosaurs, including the
earliest dinosaurs.
Triassic
continental sediments that contain vertebrate fossils are rare, thus the
importance of the Ischigualasto Basin, which contains a continuous sequence
of fossil-bearing continental sediments from the Middle Triassic Chañares
Formation and the Los Rastros Formation, through the Upper Triassic
Ischigualasto and Los Colorados.
In 1958 a
joint expedition from Harvard University and the University of La Plata
directed by Alfred S. Romer and Rosendo Pascual, discovered the highly
fossiliferous nature of the Ischigualasto Basin. Subsequent exploration and
research by the University of Tucumán, University of Chicago and the
University of San Juan has produced hundreds of specimens. The Ischigualasto
Basin documents, especially the changes in evolution and faunal replacement
during the Middle Triassic to the end of the Late Triassic, and probably the
transition to the Lower Jurassic.
A diverse
assemblage of fossil vertebrates has been discovered in the Ischigualasto
Basin. The fauna includes a very unique “transition fauna” from the Middle
Triassic Chañares Formation, a diverse Carnian fauna which includes the most
primitive and oldest dinosaurs from the Ischigualasto Formation (228 m.y).
At the top is the Los Colorados Formation which documents the latest
Triassic faunal assemblage and probably the topmost level in the transition
to Lower Jurassic faunas.
In addition
to scientific interest, there has been a popular and governmental response
to the discoveries. Ischigualasto is a harsh but beautiful valley, nicknamed
Valley of the Moon, consisting of white and variegated sandstone and
claystone, dominated by 500 foot high red cliffs. The total extent is about
5 miles wide and 40 miles long.
In 1941,
the provincial government of San Juan declared the area a Provincial Park
and designated The Museum of Natural History and the University of San Juan
as the controlling institutions for research and custody of fossil material.
The park is a popular tourist site as well as a principal area of
paleontological research. So critical is Ischigualasto to our understanding
of archosaur and mammal evolution that this park not only has gained
national protection but also has recently been recognized as a World
Heritage Site by UNESCO.
Six years
of Earthwatch supported fieldwork has significantly increased the number of
important fossils recovered from the Triassic of Ischigualasto, including a
new species of a gracile theropod closely related to the primitive
Eoraptor, new specimens of primitive dinosaurs (Herrerasaurus and
Eoraptor), the first discovery of infant rhynchosaurs grouped
together, two new species of primitive archosauriphorms closely related to
the Middle Triassic Chanaresuchus, the first record of postcraneal
remains of the proterochampsid Proterochampsa, two new specimens of
the rare poposaurid Sillosuchus, a skull of a new carnivorous
cynodont and several partial skeletons of aetosaurs.
All this
findings are contributing to better understand the evolution of these groups
and complete our picture of the biodiversity during the Triassic. Graduate
students have realized a big part of the descriptions, analysis and
interpretation as part of Masters and PhD thesis, so this project has a
great impact in education and professional training.
Besides the
discoveries of new species or new elements of known species, we collected
one of the most complete collections in the world of rynchosaurs and
cynodonts. Many of these specimens are complete and exquisite preserved
skeletons, they are already part of the permanent exhibit of the Museo de
Ciencias Naturales in San Juan and also part of traveling exhibits.
The
Ischigualasto Basin is huge and extremely rich, and the fast erosion
provides hundred of new specimens every year, so our work in these outcrops
will never end.
2.
Research Objectives
•
Continuous exploration for
new species and better-preserved specimens of known forms in the south part
of the Ischigualasto Basin near
“Agua de la Peña” where the classic locations of the Carnian Ischigualasto
Formation are (see map). We will prospect carefully the bottom of levees and
crevasse splay channels looking for transported small vertebrates remains
and microfossils. We will also prospect the northern outcrops of this
Formation near to “La Chilca” (see map). This area has been unprospected
for more than 30 years.
•
Exploration for paleovertebrates
in the Los Colorados Formation, especially the middle and upper part of the
section. We will prospect with more detail the reduced areas of overbanks
deposits, which are in the south of the basin and the north areas known as
“Los Jachaleros” and “La Esquina” (see map). The upper part of the formation
will be followed laterally in all its extension south-north to determine the
existence of Lower Jurassic sediments laying on the top of the Los Colorados
Formation. There is no clear information about the exact location of many
taxas collected during the last 20 years in the top of the Formation, but
the faunal assemblage shows a mixture of Upper Triassic and Lower Jurassic
association.
•
Careful exploration of the
transition zone between Ischigualasto and Los Colorados Formation in “La
Chilca” area (see map). This is the only part of the basin where
paleovertebrates remains were sited in the transition zone. The potential
faunal assemblage of this part of the section will give us very important
information about the faunal replacement and possible extinctions between
the Carnian and Norian.
•
Gathering sediments for washing
and acid treatment as part of the search for microfossils (pieces and
fragments of very small individuals of any taxa). We hope to find mammal
teeth or small fragments of lizards or amphibians. It is important to note
that until recently, no microvertebrates were found in the Ischigualasto
Basin. For this reason, we will try exploring different kinds of layers this
season and will treat sediments from the Los Colorados Formation for the
first time.
•
Detailed description of fossils
in situ and their relationship to the sediments and structures of the
layer in which they were found. The description of the skeletons should
include, detailed drawings of the specimen, taxa identification, type of
preservation, parts of the skeleton found, examination for signs of
transportation or action of predators, and description of the surface of the
bones to detect weathering before burial. (If the remains where transported
we will measure the direction of the current.) The architectural arrangement
of the layers where the fossil were found will be described, photographed
and mapped.
•
Detailed mapping of all
the paleovertebrate remains found using aerial photographs (scale 1:500) and
transparencies. We will use GPS to position all the collected remains.
•
Exploration for more volcanic
ash and basalt layers in the Ischigualasto and Los Colorados Formation. The
ashes will be dated using Ar/Ar and the basalts using U/Pb. At present, we
have just one trustworthy radimetric date from the bottom of the
Ischigualasto Formation, which gives us an age of 228 million years. The
exact age for a different part of the section of the Ischigualasto Basin
will allow us to determine rates of depositions, global biostratigraphy and
correlations with Triassic faunal assemblages from Brazil, USA, China,
Europe and Africa.
•
Detailed documentation and
sampling of distribution, nature, architecture, and carbonates of paleosols
in the Ischigualasto Formation. Documentation of occurrence and distribution
of plant megafossils in the Ischigualasto Formation. Lab analysis of
paleosol carbonates to be completed after the field season. All of the data
will be used for the paleoclimatic interpretation of the Ischigualasto
Formation.
3.
Methods
Our methods
are those standard for geology and paleontology. Geologists will do detailed
vertical cross sections using a Jacob Stake, a Brunton compass, a geologic
hammer, geologic magnifier glasses and a waterproof field book. We will map
on base maps and aerial photos and GPS.
Fossils,
when discovered by volunteers, will be marked with toilet paper and mapped
using a GPS. PIs will check all the findings at the end of the day or the
next day. PIs will determine the finding’s importance, and later an
extraction list will be made. The fossils to be recovered will be uncovered
using brushes and pick or dental tools, then sealed with hardener (lacquer
or very thin superglue). Then, they will be protected with aluminum foil and
covered with plaster jackets, labeled and catalogued. Most of the time, the
extraction will involve the use of heavy hammers, chisels and a power cut
saw (saw is not to be operated by volunteers).
4.
Application of Results
Vertebrate
Paleontology and probably Paleobotany will be the principal beneficiaries of
the results of our research. Sedimentary geology and basin analysis will
also benefit from our work.
Our work
has had a positive effect on public policy with regard to preservation of
fossil sites. Ischigualasto is the first public park in Argentina to be
created on the basis of its paleontological importance. The publicity
resulting from discoveries made here has had a significant effect on the
public’s awareness of fossils.
The Museum
has an active School Service division, which will benefit directly from our
field work. The increase in specimens makes it possible to loan material to
other institutions for display and to make a traveling exhibition to take to
schools.
4.1. Scientific
Contributions and Thesis
4.1.1 List
of papers and presentations thanks to Earthwatch support:
1995 Alcober O., Martínez R. y Milana J.
P.
Paleoecología de la formación Ischigualasto.
XII Jornadas Argentinas
de Paleontología de Vertebrados. Tucumán.
1995 Milana J. P.,
Alcober O. A. y Martínez R.
La mega-arquitectura
depositacional de la Formación Ischigualasto como control paleoecológico de
las faunas de paleovertebrados.
XII Jornadas Argentinas
de Paleontología de Vertebrados. Tucumán.
1995 Alcober O. A., Martínez R., Milana
J. P., Bianchi J. and Bauzá L.
A new fossil locality from the Cretaceous of
Mendoza, Argentina.
Abstracts of Papers, 55th Annual Meeting
Society of Vertebrate Paleontology. Pittsburgh.
1995 Milana J. P., Martínez R. and
Alcober O. A.
A new dinosaur locality of Upper
Triassic-Lower Jurassic age, from San Juan province, Argentina.
Abstracts, 55th Annual Meeting Society of
Vertebrate Paleontology, Pittsburgh.
1995 Martínez R., Milana, J. P. and
Alcober O. A.
Taphonomy of a basal portion of the
Ischigualasto Formation, Upper Triassic, San Juan province, Argentina.
Abstracts of Papers, 55th Annual Meeting
Society of Vertebrate Paleontology. Pittsburgh.
1996 Martínez, R.N., Forster, C.A. and
May, C.L.
The skull of Probelesodon sanjuanensis, sp. nov., from the Late
Triassic Ischigualasto Formation of Argentina.
Journal of Vertebrate Paleontology 16(2):285-291
1996 Martínez, R.N.and Forster, C.A.
A new carnivorous cynodont from the
Ischigualasto Formation (Late Triassic, Argentina) with comments on
eucynodont phylogeny.
Journal of Vertebrate Paleontology 16(2):271-284
1997 Alcober, O. A. and Parrish, M.
A new Poposaurid from the Upper Triassic of
Argentina.
Journal of Vertebrate Paleontology.
18, 2.
1997 Alcober, Oscar;
Juan P. Milana, Ricardo Martínez y Judith Parrish
Evidencias del cambio
climático global Triásico en el sur sudamericano
III Reunión Argentina del
Triásico. La Rioja, Argentina.
1997 Parrish, Judith; Oscar Alcober, Juan
Milana and Ricardo Martínez.
The Pangean Monsoon Maximum in South and
North America
GSA Meeting.
Salt Lake City, USA.
1999 Kocogian C.,
Spalleti J., Artabe A., Alcober O., Martínez R., Milana J. y Zamuner A.
Cuencas Triásicas del
Noroeste Argentino.
In:
Geología Regional Argentina, Tomo 2.
Buenos Aires.
2000 Alcober, Oscar
Redescription of the skull of Saurosuchus
galilei (Archosauria: Rauisuchidae)
Journal of Vertebrate Paleontology. 20, 2.
Sereno P., Martínez
R. and Alcober O. A.
The primitive dinosaur
Eoraptor lunensis.
In press: Special
Papers of the Society of Vertebrate Paleontology.
4.1.2. Ph.D.
and Master’s Thesis supported by Earthwatch:
1994 Martínez, R. N.
Estratigrafía del Sector
Agua de la Peña – Río de la Chilca, Formación Ischigualasto (Triásico
Superior), Cuenca de Ischigualasto, San Juan.
Master’s Thesis,
Universidad Nacional de San Juan. 210p.
1995 López, Sergio
Estudio Estratigrafico
Secuencial y Oleogeoquimico del paleolago Ischichuca-Los Rastros en el
Parque Ischigualasto, comparación con los sistemas de Mendoza Norte, y su
relación con la exploración de Hidrocarburos.
Master Thesis,
Universidad Nacional de San Juan. 187p.
1996 Alcober, O. A.
Revisión de los
Crurotarsi (Arcosauria). Tafonomía de la Formación Ischigualasto (Triásico
Superior).
Ph.D. Thesis, Universidad
Nacional de San Juan. 265p.
4.1.3. Ph.D. and Master’s Thesis in progress supported
by Earthwatch:
____ Colombi, Carina
Arquitectura aluvial,
tafonomía y distribución de paleovertebrados en un sector basal de la
Formación Ischigualasto (Triásico Superior –Cuenca de Ischigualasto).
Master Thesis,
Universidad Nacional de San Juan
____ Heredia, Guillermo
Filogenia de
Arcosauriformes (Amniota – Reptilia)
Ph D Thesis, Universidad
Nacional de San Juan
5.
Field Training
The following topics will
be covered:
·
Mapping on air photos and base
map
·
Basic anatomy of tetrapods,
especially basic anatomic features to be recognized in the specimens
(general archosaurs morphology, dinosaurs, cynodonts, mammals and
rynchosaurs).
·
Different techniques to prospect
and find fossils. How to differentiate fossils from concretions. How to mark
and catalogue fossils. How to protect and recover vertebrate fossils.
·
Recognition of geologic
features, such as paleosols, paleochannels, faults. Differences between
alluvial and lacustrine environments. Sedimentary structures such as cross
beddings, concretions, patches, roots, etc.
·
Basic introduction to archosaurs
and synapsids phylogeny and taxonomy.
·
How to use a GPS and compass.
6.
Volunteer Assignments
Volunteers will be
completely integrated into the project, doing the same tasks as the
professional staff members, perhaps at a more basic level. They will always
be with a staff member. Work parties will consist of 2, 3 or 4 people. Teams
will prospect for fossils, mark locations, map localities, gather and wash
sediments, excavate and plaster specimens. Part of the team will backpack
into areas of difficult access for preliminary exploration of new areas. In
the evening everyone will gather and chop firewood, cook and clean up, after
which we will have discussions, write in the field log, classify and catalog
specimens. We strongly recommend that volunteers have previous camping
experience. Knowledge of Spanish is useful but not essential.
7.
Project Staff
Principal
Investigators
Oscar Alcober,
Ph.D., age 39.
2001-present, Full Professor and Director of
the Museo de Ciencias Naturales, Universidad Nacional de San Juan;
1998-2000, Associate Professor and Curator
Universidad Nacional de San Juan;
1998-2000, Research Associated and collections
manager of the Texas Memorial Museum;
1997-1998, Post-doc. Visiting Scholar
University of Texas;
1996-1997, Prof. of Computer Design Univ. Nac.
of San Juan;
1994-1997, Prof. of Paleontology “Profesorado
de Biología”;
1995, Universidad Nacional de San Juan;
1994-1997, Curatorial Assoc.
Museo de Ciencias Naturales.
Ricardo Martínez,
Mr.D., age 47.
2001-present, Associate
Professor and Curator Universidad Nacional de San Juan;
1997-present, Collections
Manager Museo de Ciencias Naturales;
1995 –2000, Curatorial Assoc.
Museo de Ciencias Naturales;
1994, Universidad Nacional
de San Juan.
Guillermo Heredia,
Mr.D., age 30.
1998–present, Curatorial Assoc.
Museo de Ciencias Naturales;
1999-2000, Assistance
Professor Universidad Nacional de San Juan;
1997-2000, Prof. of
Comparative Anatomy in the “Profesorado de Biología”;
1997, Universidad Nacional
de San Luis.
Staff
Assistants
Raúl Gordillo, age
29, lab and field technician.
Diego Abelín, age 24, lab and field technician
Jimena Trotteyn, age 26, lab and field technician |